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Chapter 18

Evolution and the Origin of Species

Book Version 32
By Boundless
Boundless Biology
Biology
by Boundless
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Section 1
Understanding Evolution
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What is Evolution?

Evolution, the unifying theory of biology, describes a mechanism for the change and diversification of species over time.

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Charles Darwin and Natural Selection

Charles Darwin and Alfred Wallace independently developed the theories of evolution and its main operating principle: natural selection.

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The Galapagos Finches and Natural Selection

The differences in shape and size of beaks in Darwin's finches illustrate ongoing evolutionary change.

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Processes and Patterns of Evolution

Natural selection can only occur in the presence of genetic variation; environmental conditions determine which traits are selected.

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Evidence of Evolution

Evidence for evolution has been obtained through fossil records, embryology, geography, and molecular biology.

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Misconceptions of Evolution

There are many misconceptions about evolution, including the meaning of the word theory, the way populations change, and the origin of life.

Section 2
Formation of New Species
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The Biological Species Concept

A species is defined as a group of individuals that, in nature, are able to mate and produce viable, fertile offspring.

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Reproductive Isolation

Reproductive isolation, through mechanical, behavioral, and physiological barriers, is an important component of speciation.

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Speciation

Speciation is an event in which a single species may branch to form two or more new species.

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Allopatric Speciation

Allopatric speciation occurs when a single species becomes geographically separated; each group evolves new and distinctive traits.

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Sympatric Speciation

Sympatric speciation occurs when two individual populations diverge from an ancestral species without being separated geographically.

Section 3
Hybrid Zones and Rates of Speciation
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Hybrid Zones

Over time, two species may further diverge or reconnect, depending on the fitness strength and the reproductive barriers of the hybrids.

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Varying Rates of Speciation

Two patterns are currently observed in the rates of speciation: gradual speciation and punctuated equilibrium.

Section 4
Evolution of Genomes
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Genomic Similiarities between Distant Species

Genomic similarities between distant species can be established via analysis of genomes using advanced technology.

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Genome Evolution

Processes such as mutations, duplications, exon shuffling, transposable elements and pseudogenes have contributed to genomic evolution.

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Whole-Genome Duplication

Whole-genome duplication is characterized by an organisms entire genetic information being copied once or multiple times.

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Gene Duplications and Divergence

Gene duplications create genetic redudancy and can have various effects, including detrimental mutations or divergent evolution.

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Noncoding DNA

Noncoding DNA are sequences of DNA that do not encode protein sequences but can be transcribed to produce important regulatory molecules.

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Variations in Size and Number of Genes

The genome size does not always correlate with the complexity of the organism and, in fact, shows great variation in size and gene number.

Section 5
Evidence of Evolution
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The Fossil Record as Evidence for Evolution

Fossils tell us when organisms lived, as well as provide evidence for the progression and evolution of life on earth over millions of years.

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Fossil Formation

Fossils can form under ideal conditions by preservation, permineralization, molding (casting), replacement, or compression.

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Gaps in the Fossil Record

Because not all animals have bodies which fossilize easily, the fossil record is considered incomplete.

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Carbon Dating and Estimating Fossil Age

The age of fossils can be determined using stratigraphy, biostratigraphy, and radiocarbon dating.

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The Fossil Record and the Evolution of the Modern Horse

The detailed fossil record of horses has provided insight into their evolutionary progress.

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Homologous Structures

Homologous structures are similar structures that evolved from a common ancestor.

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Convergent Evolution

Convergent evolution occurs in different species that have evolved similar traits independently of each other.

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Vestigial Structures

Vestigial structures have no function but may still be inherited to maintain fitness.

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Biogeography and the Distribution of Species

The biological distribution of species is based on the movement of tectonic plates over a period of time.

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Evolution and the Origin of Species
  • Understanding Evolution
  • Formation of New Species
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The Evolution of Populations
  • Population Evolution
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